Gamera's size rivals that of his famous daikaiju counterpart, Godzilla. Unlike any other species of turtles, Gamera has the habit of walking bipedal rather than all fours, though he occasionally walks like a quadruped in his first three films. Gamera was capable of using his upper limbs in the same manner as Godzilla, as his forelegs had appendages much closer in construction to hands than feet, and was capable of grappling with opponents and manipulating objects. His mouth is filled with teeth, unlike any living modern turtle (several types of extinct prehistoric turtles were toothed, however), with a pair of large tusks protruding upward from his lower jaw. Gamera is also usually seen with very large human-like eyes, adding intelligence to his overall appearance.